Recent Publications


  • Burks, R.A., L. Masner, N.F. Johnson and A.D. Austin. Systematics of the parasitic wasp genus Oxyscelio Kieffer (Hymenotpera: Platygastridae s.l.), part III: African fauna. Submitted for publication to ZooKeys, 11 Nov 2015.
  • Azevedo, C.O., A. Dal Molin, A. Penteado-Dias, A.C.C. Macedo, B. Rodriguez-V., B.Z.K. Dias, C. Waichert, D. Aquino, D.R. Smith, E.M. Shimbori, F.B. Noll, G. Gibson, H.C. Onody, J.M. Carpenter, J.E. Lattke, K. dos S. Ramos, K. Williams, L. Masner, L.S. Kimsey, M.T. Tavares, M. Olmi, M.L. Buffington, M. Ohl, M. Sharkey, N.F. Johnson, R. Kawada, R.B. Gonçalves, R.M. Feitosa, S. Heydon, T.M. Guerra, T.S.R. da Silva and V. Costa. 2015. Checklist of the genera of Hymenoptera (Insecta) from Espírito Santo state, Brazil. Boletim do Museum Biológico Mello Leitão (n.s.) 37:313–343.
  • Talamas, E.J., N.F. Johnson and M. Buffington. 2015. Key to Nearctic species of Trissolcus (Hymenoptera, Scelionidae), natural enemies of native and invasive stink bugs (Hemiptera, Pentatomidae). Journal of Hymenoptera Research 43:45-110. doi:10.3897/JHR.43.8560. PDF available


  • Matsuo, K., Y. Hirose and N.F. Johnson. 2014. A taxonomic issue of two species of Trissolcus (Hymenoptera: Platygastridae) parasitic on eggs of the brown-winged green bug, Plautia stali (Hemiptera: Pentatomidae): resurrection of T. plautiae, a cryptic species of T. japonicus revealed by morphology, reproductive isolation and molecular evidence. Applied Entomology and Zoology. doi:10.1007/s13355-014-0260-4. PDF available
  • van Noort, S., L. Masner, O. Popovici, A.A. Valerio, C. Taekul, N.F. Johnson, N.P. Murphy and A.D. Austin. 2014. Systematics and biology of the aberrant intertidal parasitoid wasp Echthrodesis lamorali Masner (Hymenoptera, Platygastridae s.l.): a parasitoid of spider eggs. Invertebrate Systematics 28: 1-16.
  • Mao, M., A.D. Austin, N.F. Johnson and M. Dowton. 2014. Coexistence of minicircular and a highly rearranged mtDNA molecule suggests that recombination shapes mitochondrial genome organization. Molecular Biology and Evolution 31: 636-644.
  • Yoder, M.J., A.A. Valerio, A. Polaszek, S. van Noort, L. Masner and N.F. Johnson. 2014. Monograph of the Afrotropical species of Scelio Latreille (Hymenoptera: Platygastridae), egg parasitoids of acridid grasshoppers (Orthoptera: Acrididae). ZooKeys 380: 1-188.PDF available


  • Johnson, N.F., R. Burks, A. Austin and Z.-F. Xu. 2013. Chinese species of egg-parasitoids of the genera Oxyscelio Kieffer, Heptascelio Kieffer and Platyscelio Kieffer (Hymenoptera: Platygastridae s.l., Scelioninae). Biodiversity Data Journal doi:10.3897/BDJ.1.e987 PDF available
  • Burks, R.A., L. Masner, N.F. Johnson and A.D. Austin. 2013. Systematics of the parasitic wasp genus Oxyscelio Kieffer (Hymenoptera, Platygastridae s.l.), part II: the Australian and southwest Pacific fauna. ZooKeys 331: 1-266.PDF available
  • Taekul, C., A.A. Valerio, A.D. Austin, H. Klompen and N.F. Johnson. 2013. Molecular phylogeny of telenomine egg parasitoids (Hymenoptera: Platygastridae s.l.: Telenominae): evolution of host shifts and implications for classification. Systematic Entomology doi:10.1111/syen.12032PDF available
  • Aguiar, A. P., A. R. Deans, M. S. Engel, M. Forshage, J. T. Huber, J. T. Jennings, N. F. Johnson, A. S. Lelej, J. T. Longino, V. Lohrmann, I. Mikó, M. Ohl, C. Rasmussen, A. Taeger and D. S. K. Yu. 2013. Order Hymenoptera. Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa 3703: 51-62.PDF available
  • Talamas, E.J., L. Masner and N.F. Johnson. 2013. Systematics of Trichoteleia Kieffer and Paridris Kieffer (Hymenoptera: Platygastroidea, Platygastridae). Journal of Hymenoptera Research 34:1-79.PDF available
  • Valerio, A.A., A.D. Austin, L. Masner and N.F. Johnson. 2013. Systematics of Old World Odontacolus Kieffer s.l. (Hymenoptera, Platygastridae, s.l.): parasitoids of spider eggs. ZooKeys 314:1-151.PDF available
  • Chen, H.-Y., N.F. Johnson, L. Masner and Z.-F. Xu. The genus Macroteleia Westwood (Hymenoptera, Platygastridae, Scelioninae) from China. ZooKeys 300:1-98.PDF available
  • Burks, R.A., L. Masner, N.F. Johnson and A.D. Austin. 2013. Systematics of the parasitic wasp genus Oxyscelio Kieffer (Hymenoptera, Platygastridae s.l.), part I: Indo-Malayan and Palearctic fauna. ZooKeys 292:1-263.PDF available
  • Johnson, N.F. 2013. Hymenoptera. Pages 177-184, in S.A. Levin, ed., Encyclopedia of Biodiversity, second edition. Vol. 4. Academic Press, Waltham, MA.


  • Burks, R.A., L. Masner, N.F. Johnson and A.D. Austin. 2012. Taxonomic revision of Bracalba Dodd (Hymenoptera: Platygastridae s.l.), a parasitoid wasp genus endemic to Australia. ZooKeys 236:1–53.PDF available
  • Singh, L.R.K., N.F. Johnson and V.V. Ramamurthy. 2012. New species of the genus Psix Kozlov & Lê (Hymenoptera: Platygastridae). Zootaxa 3530:35–42.

2012, continued

  • Talamas, E.J., L. Masner and N.F. Johnson. 2012. Paridris Kieffer of the New World (Hymenoptera: Platygastroidea, Platygastridae). ZooKeys 233:31–90.PDF available
  • Popovici, O.A. and N.F. Johnson. 2012. Gross anatomy of the Malpighian tubulues and internal male genitalia of Scelioninae (Hymenoptera; Platygastroidea; Platygastridae) with phylogenetic implications. Proceedings of the Entomological Society of Washington 114:372–397.
  • Seltman, K.C., M.J. Yoder, I. Mikó, M. Forshage, M.A. Bertone, D. Agosti, A.D. Austin, J.P. Balhoff, M.L. Borowiec, S.G. Brady, G.R. Broad, D.J. Brothers, R.A. Burks, M.L. Buffington, H.M. Campbell, K.J. Dew, A.F. Ernst, J.L. Fernández-Triana, M.W. Gates, G.A.P. Gibson, J.T. Jennings, N.F. Johnson, D. Karlsson, R. Kawada, L. Krogmann, R.R. Kula, P.L. Mullins, M. Ohl, C. Rasmussen, F. Ronquist, S. Schulmeister, M.J. Sharkey, E. Talamas, E. Tucker, L. Vilhelmsen, P.S. Ward, R.A. Wharton and A.R. Deans. 2012. A hymenopterist’s guide to the Hymenoptera Anatomy Ontology: utility, clarification, and future directions. Journal of Hymenoptera Research 27:67–88.
  • Mao, M., A. Valerio, A.D. Austin, M. Dowton and N.F. Johnson. 2012. The first mitochondrial genome for the wasp superfamily Platygastroidea: the egg parasitoid Trissolcus basalis. Genome 55: 194-204.PDF available
  • Johnson, N.F. 2012. A collaborative, integrated and electronic future for taxonomy. Invertebrate Systematics 25: 471–475.PDF available
  • Johnson, N.F. and L. Musetti. 2012. Genera of the parasitoid wasp family Monomachidae (Hymenoptera: Diaprioidea). Zootaxa 3188: 31–41.PDF available


  • Johnson, N.F. and L. Musetti. 2011. Redescription and revision of the Neotropical genus Pseudoheptascelio Szabó (Hymenoptera, Platygastridae, Scelioninae), parasitoids of eggs of short-horned grasshoppers (Orthoptera, Acrididae). ZooKeys 136:93–112.PDF available
  • Talamas, E., L. Masner and N.F. Johnson. 2011. Revision of the Paridris nephta species group (Hymenoptera, Platygastroidea, Platygastridae). ZooKeys 133:49–94.PDF available
  • Talamas, E., L. Masner and N.F. Johnson. 2011. Revision of the Malagasy genus Trichoteleia Kieffer (Hymenoptera, Platygastroidea, Platygastridae). ZooKeys 80:1–126.PDF available
  • Triplehorn, C.A. and N.F. Johnson. 2011. Estudo dos insectos. Tradução da 7ª edição de Borror and Delong’s introduction to the study of insects. Cengage Learning, São Paulo. 809 pp.


  • Penev, L., D. Agosti, T. Georgiev, T. Catapano, J. Miller, V. Blagoderov, D. Roberts, V.S. Smith, I. Brake, S. Ryrcroft, B. Scott, N.F. Johnson, R.A. Morris, G. Sautter, V. Chavan, T. Robertson, D. Remsen, P. Stoev, C. Parr, S. Knapp, W.J. Kress, F.C. Thompson, and T. Erwin. 2010. Semantic tagging of and semantic enhacements to systematics papers: ZooKeys working examples. ZooKeys 50:1–16.
  • Taekul, C., N.F. Johnson, L. Masner, A. Polaszek and Rajmohana K. 2010. World species of the genus Platyscelio Kieffer (Hymenoptera, Platygastridae). ZooKeys 50:97–126.PDF available
  • Iranipour, S. and N.F. Johnson. 2010. Trissolcus radjabii n.sp. (Hymenoptera: Platygastridae), an egg parasitoid of the shield bug, Apodiphus amygdale (Heteroptera: Pentatomidae) and the sunn pest, Eurygaster integriceps Puton (Heteroptera: Scutelleridae). Zootaxa 2515:65–68.
  • Johnson, N.F. 2010. Future taxonomy today: new tools applied to accelerate the taxonomic process. Pages 137–147, in A. Polaszek, ed., Systema Naturae 250: The Linnaean Ark. CRC Press Taylor & Francis Group, London. 276 pp.